Santiago Fraga



Publications in the last 5 years


Kohli, G.S., Campbell, K., John, U., Smith, K.F., Fraga, S., Rhodes, L.L., Murray, S.A., 2017. Role of Modular Polyketide Synthases in the Production of Polyether Ladder Compounds in Ciguatoxin-Producing Gambierdiscus polynesiensis and G. excentricus (Dinophyceae). Journal of Eukaryotic Microbiology, n/a-n/a.

Pisapia, F., Holland, W.C., Hardison, D.R., Litaker, R.W., Fraga, S., Nishimura, T., Adachi, M., Nguyen-Ngoc, L., Séchet, V., Amzil, Z., Herrenknecht, C., Hess, P., 2017. Toxicity screening of 13 Gambierdiscus strains using neuro-2a and erythrocyte lysis bioassays. Harmful Algae 63, 173-183.

Berdalet, E., Tester, P.A., Chinain, M., Fraga, S., Lemée, R., Litaker, W., Penna, A., Usup, G., Vila, M., Zingone, A., 2017. Harmful algal blooms in benthic systems: Recent progress and future research. Oceanography 30(1), 36-45.


Fraga, S., Rodríguez, F., Riobó, P., Bravo, I., 2016. Gambierdiscus balechii sp. nov (Dinophyceae), a new benthic toxic dinoflagellate from the Celebes Sea (SW Pacific Ocean). Harmful Algae, 58, 93-105.

Delgado, G., Arencibia, G., Lucas, F., Rosquete, C.M., Fraga, S., 2016. A Cochlodinium polykrikoides Margalef bloom in the channels of Marina Hemingway, La Habana, Cuba. Harmful Algae News 53, 8.
Tester, P.A., Fraga, S. Regional Training Course on Ciguatera Fish Poisoning (CFP): Field Monitoring and Toxin Data Management. Santo Domingo, October 26 – November 6, 2015. Harmful Algae News 53:19-20.

Álvarez, G., Uribe, E., Regueiro, J., Blanco, J., Fraga, S., 2016. Gonyaulax taylorii, a new yessotoxins-producer dinoflagellate species from Chilean waters. Harmful Algae 58, 8-15.


Fraga, S., Sampedro, N., Larsen, J., Moestrup, O., Calado, A.J., 2015. Arguments against the proposal 2302 by John & al. to reject the name Gonyaulax catenella (Alexandrium catenella). Taxon 64(3), 634-635.
Nascimento, S.M., Melo, G., Salgueiro, F., Diniz, B.d.S., Fraga, S., 2015. Morphology of Gambierdiscus excentricus (Dinophyceae) with emphasis on sulcal plates. Phycologia 54(6), 628-639.

Penna, A., Perini, F., Dell’Aversano, C., Capellacci, S., Tartaglione, L., Giacobbe, M.G., Casabianca, S., Fraga, S., Ciminiello, P., Scardi, M., 2015. The sxt Gene and Paralytic Shellfish Poisoning Toxins as Markers for the Monitoring of Toxic Alexandrium Species Blooms. Environmental Science & Technology 49(24), 14230-14238.

Pitcher, G.C., Fraga, S., 2015. Harmful algal bloom events in the Canary Current Large Marine Ecosystem, In: Valdés, L., Déniz‐González, I. (Eds.), Oceanographic and biological features in the Canary Current Large Marine Ecosystem. IOC‐UNESCO, Paris, pp. 175‐182.


Penna, A., Battocchi, C., Capellacci, S., Fraga, S., Aligizaki, K., Lemée, R., Vernesi, C. 2014. Mitochondrial, but not rDNA, genes fail to discriminate dinoflagellate species in the genus Ostreopsis. Harmful Algae 40:40-50.

Fraga, S., Rodríguez, F. 2014. Genus Gambierdiscus in the Canary Islands (NE Atlantic Ocean) with Description of Gambierdiscus silvae sp. nov., a New Potentially Toxic Epiphytic Benthic Dinoflagellate. Protist 165:839-853.

Bravo, I., Figueroa, R.I., Fraga, S. 2014. Cellular and nuclear morphological variability within a single species of the toxigenic dinoflagellate genus Gambierdiscus: Relationship to life-cycle processes. Harmful Algae 40:1-8.

Figueroa, R.I., Cuadrado, A., Stüken, A., Rodríguez, F., Fraga, S. 2014. Ribosomal DNA Organization Patterns within the Dinoflagellate Genus Alexandrium as revealed by FISH: Life Cycle and Evolutionary Implications. Protist, 165:343-363.


Amorim, A., Veloso, V., Rodríguez, F. & Fraga, S. 2013. Life-cycle, morphology and phylogeny of species of Fragilidium Balech from west Iberia In: Lewis, J. M., Marret, F. & Bradley, L. [Eds.] Bilogical and Geological Perspectives of Dinoflagellates. The Micropaleontological Society, London, pp. 223–32.

Mohammad-Noor, N., Moestrup, Ø., Lundholm, N., Fraga, S., Adam, A., Holmes, M. J. & Saleh, E. 2013. Autecology and phylogeny of Coolia tropicalis and Coolia malayensis (Dinophyceae), with emphasis on taxonomy of Coolia tropicalis based on light microscopy, scanning electron microscopy and LSU rDNA. Journal of Phycology 49:536–45.